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Season five of Murdoch Mysteries opens in unfamiliar territory, as Detective William Murdoch (Yannick Bisson) leaves Toronto to try his luck while the Klondike Gold Rush in the Yukon winds down. Suspended for interfering in an investigation, on the run for surreptitiously freeing a murderer to make amends, and nursing a broken heart after Dr. Ogden's marriage to another man, he tries to escape his past, and his passion for detective work. Back in the city, newly promoted Chief Constable Giles (Nigel Bennett) makes it clear to Inspector Thomas Brackenreid (Thomas Craig), that he'll be keeping a very close eye on police matters. Meanwhile, Murdoch's beloved former associate, Dr. Julia Ogden (Hélène Joy), is now working alongside Acting Detective George Crabtree (Jonny Harris) and Dr. Emily Grace (Georgina Reilly), a feisty young protégé. In the remote north, Murdoch is pleased to discover that his fledgling prospecting career may garner great success, a small comfort after Julia's marriage to another man. But as he tries to rebuild his life, he finds himself drawn into the case of Mrs. Elizabeth Bryant (Jill Halfpenny), an innkeeper charged with murder. The plot thickens when writer Jack London (Aaron Ashmore) lends his insights, and Murdoch soon realizes that while he had hoped to leave his shattered heart and tattered career back in Toronto, his days as a detective are not behind him.
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Murdoch is in Bristol, England, with no memory of who he is and two gunmen chasing him through the streets. With the help of a local pub Landlady he starts to piece together an assassination plot. In Toronto Detective Hamish Sloack is assigned to help find the missing Murdoch.
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Murdoch and Inspector Brackenreid are aboard the SS Keewatin after the owner, Clarence MacFarlane, receives what he believes is a threat. The Keewatin is about to depart from Toronto on its maiden voyage destined for Rochester, N.Y. and Murdoch thinks they should stay on board, especially after he sees that Julia is one of the invited guests. Their presence proves useful after MacFarlane's daughter Amy, who is engaged to her father's chief financial officer Owen Mathers, falls overboard and is nowhere to be found. As Murdoch investigates he finds there is a real threat against the Keewatin and the true facts behind Amy's disappearance.
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第四季里茱蒂和贝蒂吵架，去了纽约。本季中贝蒂为求得她的原谅而精心准备，却事与愿违。海伦娜因为偷钱获刑入狱。在狱中海伦娜遭到监狱犯人的骚扰，幸好得到狱友——黑人女孩的解救，两人坠入爱河。詹妮变得比以前更加神经质跟冷漠，她仗着有投资人撑腰，接替了蒂娜的位置，成为《Lez Girls》的负责人。在片场她邂逅了女主角妮基（凯特·弗兰茨 Kate French饰），两人相爱。在“星球”餐厅，詹妮的女粉丝阿黛尔（马来亚里·韦拉·德鲁 Malaya Rivera Drew饰）登场。塔莎因为自己同性恋的行为被调查，没去成伊拉克。夏恩仍然改不了沾花惹草的老毛病，最为讽刺的是，居然和妮基好上了，这让詹妮伤透了心。虽然《Lez Girls》的拍摄波折不断，最终还是完成了制作。庆功会上，贝蒂和蒂娜公然复合。
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根据畅销小说《Women of the other world》改编。主人公Elena Michaels是一名女摄影师。她一直隐藏着自己的真实身份并努力过着平常人的生活。可惜事与愿违，一通神秘来电打破了她平静的生活。从古至今一直是狼人庇护所的Stonehaven接连出现尸体，狼人族群收到未知猎人的匿名威胁。被迫回到Stonehaven的Elena能否揭开血案后的真相？她又能否从狼人的世界中全身而退？狼人族群的未来又何去何从？一切的一切都把握在世界上唯一硕果仅存的女狼人Elena的手中。（文：HTC@深影字幕组）
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根据畅销小说《Women meijubar.net of the other world》改编。主人公Elena Michaels是一名女摄影师。她一直隐藏着自己的真实身份并努力过着平常人的生活。可惜事与愿违，一通神秘来电打破了她平静的生活。从古至今一直是狼人庇护所的Stonehaven接连出现尸体，狼人族群收到未知猎人的匿名威胁。被迫回到Stonehaven的Elena能否揭开血案后的真相？她又能否从狼人的世界中全身而退？狼人族群的未来又何去何从？一切的一切都把握在世界上唯一硕果仅存的女狼人Elena的手中。